About the Program
The Department of Molecular and Cell Biology offers a program of graduate study leading to the PhD in molecular and cell biology. This program provides advanced training in the research methods and concepts of the study of the molecular structures and processes of cellular life. The training is intellectually focused, but at the same time offers an unusually wide range of opportunities for varied disciplinary specialization.
The teaching and research activities of the Department of Molecular and Cell Biology (MCB) concern the molecular structures and processes of cellular life and their roles in the function, reproduction, and development of living organisms.
This agenda covers a broad range of specialized disciplines, including biochemistry, biophysics, molecular biology, structural biology, genetics, genomics, bioinformatics, cell biology, developmental biology, tumor biology, microbiology, immunology, pathogenesis, and neurobiology.
The types of living organisms from which the departmental faculty draws its working materials are as diverse as its disciplinary specializations, ranging from viruses and microbes through plants, roundworms, annelids, arthropods, and mollusks to fish, amphibia, and mammals.
The faculty is organized into five divisions: Biochemistry, Biophysics, and Structural Biology; Cell and Developmental Biology; Genetics, Genomics and Development; Immunology and Pathogenesis and Neurobiology.
Admissions
Admission to the University
Minimum Requirements for Admission
The following minimum requirements apply to all graduate programs and will be verified by the Graduate Division:
- A bachelor’s degree or recognized equivalent from an accredited institution;
- A grade point average of B or better (3.0);
- If the applicant comes from a country or political entity (e.g., Quebec) where English is not the official language, adequate proficiency in English to do graduate work, as evidenced by a TOEFL score of at least 90 on the iBT test, 570 on the paper-and-pencil test, or an IELTS Band score of at least 7 on a 9-point scale (note that individual programs may set higher levels for any of these); and
- Sufficient undergraduate training to do graduate work in the given field.
Applicants Who Already Hold a Graduate Degree
The Graduate Council views academic degrees not as vocational training certificates, but as evidence of broad training in research methods, independent study, and articulation of learning. Therefore, applicants who already have academic graduate degrees should be able to pursue new subject matter at an advanced level without the need to enroll in a related or similar graduate program.
Programs may consider students for an additional academic master’s or professional master’s degree only if the additional degree is in a distinctly different field.
Applicants admitted to a doctoral program that requires a master’s degree to be earned at Berkeley as a prerequisite (even though the applicant already has a master’s degree from another institution in the same or a closely allied field of study) will be permitted to undertake the second master’s degree, despite the overlap in field.
The Graduate Division will admit students for a second doctoral degree only if they meet the following guidelines:
- Applicants with doctoral degrees may be admitted for an additional doctoral degree only if that degree program is in a general area of knowledge distinctly different from the field in which they earned their original degree. For example, a physics PhD could be admitted to a doctoral degree program in music or history; however, a student with a doctoral degree in mathematics would not be permitted to add a PhD in statistics.
- Applicants who hold the PhD degree may be admitted to a professional doctorate or professional master’s degree program if there is no duplication of training involved.
Applicants may apply only to one single degree program or one concurrent degree program per admission cycle.
Required Documents for Applications
- Transcripts: Applicants may upload unofficial transcripts with your application for the departmental initial review. If the applicant is admitted, then official transcripts of all college-level work will be required. Official transcripts must be in sealed envelopes as issued by the school(s) attended. If you have attended Berkeley, upload your unofficial transcript with your application for the departmental initial review. If you are admitted, an official transcript with evidence of degree conferral will not be required.
- Letters of recommendation: Applicants may request online letters of recommendation through the online application system. Hard copies of recommendation letters must be sent directly to the program, not the Graduate Division.
- Evidence of English language proficiency: All applicants from countries or political entities in which the official language is not English are required to submit official evidence of English language proficiency. This applies to applicants from Bangladesh, Burma, Nepal, India, Pakistan, Latin America, the Middle East, the People’s Republic of China, Taiwan, Japan, Korea, Southeast Asia, most European countries, and Quebec (Canada). However, applicants who, at the time of application, have already completed at least one year of full-time academic course work with grades of B or better at a US university may submit an official transcript from the US university to fulfill this requirement. The following courses will not fulfill this requirement:
- courses in English as a Second Language,
- courses conducted in a language other than English,
- courses that will be completed after the application is submitted, and
- courses of a non-academic nature.
If applicants have previously been denied admission to Berkeley on the basis of their English language proficiency, they must submit new test scores that meet the current minimum from one of the standardized tests. Official TOEFL score reports must be sent directly from Educational Test Services (ETS). The institution code for Berkeley is 4833. Official IELTS score reports must be mailed directly to our office from the British Council. TOEFL and IELTS score reports are only valid for two years.
Where to Apply
Visit the Berkeley Graduate Division application page.
Doctoral Degree Requirements
Normative Time Requirements
- Normative time to advancement is 2 years
- Normative time in candidacy is 3.5 years
- Total normative time is 5.5 years
Curriculum
| Code | Title | Units |
|---|---|---|
| MCELLBI 200A | Fundamentals of Molecular and Cell Biology | 3 |
| MCELLBI 200B | Fundamentals of Molecular and Cell Biology | 3 |
| MCELLBI 291A | Introduction to Research | 2-12 |
| MCELLBI 291B | Introduction to Research | 2-12 |
| MCELLBI 293A | Research Seminar | 2 |
| MCELLBI 293C | Responsible Conduct in Research | 1 |
| MCELLBI 293D | Rigor and Reproducibility in Research | 1 |
| MCELLBI 293R | Responsible Conduct of Research Refresher | 1 |
| MCELLBI 380 | Teaching of Molecular and Cell Biology (2 courses) | 1-2 |
| Two MCELLBI advanced topics electives | 6-8 | |
| MCELLBI 290 | Graduate Seminar (3) | 1 |
| MCELLBI 292 | Research | 3-12 |
Faculty and Instructors
+ Indicates this faculty member is the recipient of the Distinguished Teaching Award.
Faculty
Hillel Adesnik, Associate Professor. Neural basis of perception, neuroscience, neurobiology .
Research Profile
Georjana Barnes, Professor. Biochemistry, genetics, cancer, biology, microtubule cytoskeleton, cell cycle controls, cellular imaging.
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Gregory M. Barton, Professor. Immunology, cell biology, infectious disease, innate immunity.
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Helen Bateup, Assistant Professor. Molecular and cellular neuroscience, neurodevelopmental disorders, autism, epilepsy.
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Diana Bautista, Associate Professor. Ion channels, sensory physiology, chemosensation, touch, thermosensation, somatosensory system.
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Eric Betzig, Professor. Imaging microscopy, biophysics.
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David Bilder, Professor. Genetics, cancer, Drosophila, cell biology, cell polarity, tumor suppressor, epithelial architecture, polarity, and proliferation control.
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Michael R. Botchan, Professor. Eukaryotic gene expression, drosophila chromosomes, papilloma viral DNA, chromosomal dynamics.
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Gloria Brar, Assistant Professor. Meiosis, translation, sORFs, stress responses.
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Steve Brohawn, Assistant Professor. Neurobiology.
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Carlos J. Bustamante, Professor. Nanoscience, structural characterization of nucleo-protein assemblies, single molecule fluorescence microscopy, DNA-binding molecular motors, the scanning force microscope, prokaryotes.
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Jamie Cate, Professor. Molecular basis for protein synthesis by the ribosome, RNA, antibiotics, a thermophilic bacterium, escherichia coli.
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Christopher J. Chang, Professor. Chemistry, inorganic chemistry, neuroscience, bioinorganic chemistry, general physiology, organic chemistry, new chemical tools for biological imaging and proteomics, new metal complexes for energy catalysis and green chemistry, chemical biology.
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Michelle Chang, Associate Professor. Biochemistry, enzymology, synthetic biology, biophysics .
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Kathleen Collins, Professor. RNA, telomerase, Telomere function, Telomere replication.
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Jacob E. Corn, Assistant Adjunct Professor.
Laurent Coscoy, Associate Professor. Immunology, viruses, viral infection, immune responses, immune evasion.
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Jeffery S. Cox, Professor. TB, Mycobacterium tuberculosis, M tuberculosis, genetics, proteomics, transcriptional profiling, host-pathogen interactions, host-directed therapy.
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Yang Dan, Professor. Neuronal circuits, mammalian visual system, electrophysiological, psychophysical and computational techniques, visual cortical circuits, visual neurons.
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Xavier Darzacq, Associate Professor. Transcription regulation during cellular differentiation Linking the biophysical rules of nuclear organization and gene expression control mechanism .
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Karen Davies, Assistant Adjunct Professor.
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Abby Dernburg, Professor. Genomics, chromosome remodeling and reorganization during meiosis, Down syndrome, DNA.
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Andrew Dillin, Professor. Endocrinology, genetics of aging.
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Jennifer A. Doudna, Professor. RNA machines, hepatitis C virus, RNA interference, ribosomes.
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David G. Drubin, Professor. Cellular morphogenesis, plasma membrane dynamics, microtubule cytoskeletons, cytoskeletal proteins, morphological development.
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Peter H. Duesberg, Professor. Genetic structure of retroviruses, carcinogenesis, aneuploidy, virology, HIV-AIDS.
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Michael DuPage, Assistant Professor. Immunology, cancer, tumor immunity, immune responses, chronic inflammation and cancer, tumor biology, epigenetics.
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Michael B. Eisen, Professor. Genomics, genome sequencing, bioinformatics, animal development.
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+ Dan Feldman, Associate Professor. Neurobiology, learning, neurophysiology, sensory biology.
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Marla B. Feller, Professor. Neurophysiology, developmental neuroscience.
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+ Gary L. Firestone, Professor. Cancer, steroid hormones, molecular endocrinology, tumor biology, growth factors, dietary compounds, tumor cells, glucocorticoids.
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John Gerard Flannery, Professor. Neurobiology, optometry, vision science, cell and molecular biology of the retina in normal and diseased states.
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Hernan G. Garcia, Assistant Professor. Biophysics.
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Gian Garriga, Professor. Developmental neurobiology, molecular genetics, development of nervous systems, cell division, cell migration, axonal pathfinding, caenorhabditis elegans.
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Britt Glaunsinger, Associate Professor. Virology, gene expression, herpesvirus.
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Ming Chen Hammond, Assistant Professor. Molecular biology, biochemistry, organic chemistry, synthetic biology, chemical biology.
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Iswar Krishna Hariharan, Professor. Growth regulation, regeneration, cancer.
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Richard M. Harland, Professor. Molecular biology, early vertebrate development, Xenopus, embryo development.
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Lin He, Associate Professor. Comparative genomics, developmental biology, cell biology.
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Rebecca Heald, Professor. Cell division, Xenopus, mitotic spindle assembly and function, size control of intracellular structures.
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Dirk Hockemeyer, Assistant Professor. Developmental biology, cell biology.
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James Hurley, Professor. Structural Biology, reconstitution, membrane biology, autophagy, HIV, x-ray crystallography, cryo-electron microscopy.
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Nicholas Ingolia, Assistant Professor. Ribosome Profiling, translation, genomics.
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Ehud Y. Isacoff, Professor. Ion channel function, synaptic plasticity, neural excitability, synaptic transmission, the synapse.
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Na Ji, Associate Professor. Neuroscience, microscopy techniques, in vivo imaging, biophotonics, biophysics, adaptive optics, optics.
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Gary H. Karpen, Adjunct Professor. Gene expression, cell biology, chromosome structure and function, drosophila melanogaster, centromere identity and function.
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Nicole King, Professor. Genetics, developmental biology, biology, choanoflagellates, multicellularity, evolution of animals, comparative genomics, eukaryotes, host-microbe interactions, bacterial signals.
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Douglas E. Koshland, Professor. Higher order chromosome structure, genome integrity, sister chromatid cohesion, chromosome condensation, desiccation tolerance, microbial genetics.
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Richard H. Kramer, Professor. Cells, synaptic transmission, chemical signaling between neurons, ion channels, electrical signals, chemical reagents, synapses.
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John Kuriyan, Professor. Structural and functional studies of signal transduction, DNA replication, cancer therapies, phosphorylation.
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Stephan Lammel, Assistant Professor. Neuroscience, Optogenetics, dopamine, addiction, depression.
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Polina Lishko, Assistant Professor. Reproductive and Developmental Biology, ion channels, Physiology of Fertilization and Early Embryo Development.
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Ellen Lumpkin, Professor.
Kunxin Luo, Professor. Signal transduction pathways, mechanisms controlling the receptor kinases, regulation of mammary epithelial cell differentiation, breast carcinogenesis.
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Michael A. Marletta, Professor. Chemical biology, molecular biology, structure/function relationships in proteins, catalytic and biological properties of enzymes, cellular signaling, nitric oxide synthase, soluble guanylate cyclase, gas sensing, cellulose degradation, polysaccharide monooxygenases.
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Susan Marqusee, Professor. Amino acids, determinants of protein structure and folding, biophysical, structural and computational techniques, translocation, protein synthesis.
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Andreas Martin, Associate Professor. Proteasome.
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Sabeeha Merchant, Professor.
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Barbara J. Meyer, Professor. Developmental biology, gene expression, genetic determination of sex, regulatory genes, chromosome dynamics, X-chromosome.
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Craig Miller, Associate Professor. Genetics, developmental biology, evolutionary biology, evolution, quantitative genetics, developmental genetics, evolutionary genetics, craniofacial development.
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Evan W. Miller, Assistant Professor. Biochemistry, biophysics, Structural Biology .
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Priya Moorjani, Assistant Professor. Human evolutionary genetics.
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John Ngai, Professor. Nervous system, molecular and cellular mechanisms of olfaction, detection of odors, odorant receptors, olfactory neurons, DNA microarray technologies, genome-wide patterns of gene expression.
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Eva Nogales, Professor. Biochemistry, complex biological assemblies, structure and regulation of the cytoskeleton, microtubule dynamics, human transcriptional initiation machinery, biophysics.
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Daniel Normura, Associate Professor. Chemistry, molecular and cell biology, nutritional sciences and toxicology, metabolism, chemical biology, cancer, drug discovery, chemoproteomics, metabolomics.
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Eunyong Park, Assistant Professor. Protein function, Structural Biology, biochemistry, membrane proteins.
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Nipam Patel, Professor. Genetics, evolution, crustaceans, insects, arthropods, homeotic (Hox) genes, segmentation, embryonic pattern formation, neural patterning.
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Daniel A. Portnoy, Professor. Mammalian cells, molecular and cellular basis of microbial pathogenesis, defense against infection, listeria monocytogenes, cell biology of infection, mechanisms of secretion.
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Michael Rape, Professor. Cancer, protein degradation, siRNA, Berkeley Screening Center.
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David H. Raulet, Professor. Biology, pathogens, viruses, T-cell development and function, natural killer (NK) cells, lymphocyte receptors, microorganisms, cancer cells, tumor immunity.
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+ Jasper D. Rine, Professor. Biology, cell biology, DNA replication, gene regulation, saccharomyces cerevisiae, genetic analysis, genome, cholesterol biosynthetic pathway, modification of proteins, prenylated proteins.
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Donald C. Rio, Professor. Molecular genetics, drosophila melanogaster, transposable elements, RNA splicing, nucleic acid rearrangement reactions, P elements and their cellular host, HIV, proteomic diversification, nucleoprotein complexes.
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Ellen Robey, Professor. Fate determination in the T-lymphocyte lineage, T cell development in the mouse, thymic development, cellular maturation, parasitic infection, chronic infection, host-pathogen interactions, Toxoplasma gondii.
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Henk Roelink, Associate Professor. Stem cells, neural development, embryonic induction.
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Daniel S. Rokhsar, Professor. Biology, collective phenomena and ordering in condensed matter and biological systems, theoretical modeling, computational modeling, behavior of quantum fluids, cold atomic gases, high temperature superconductors, Fermi and Bose systems.
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Kaoru Saijo, Assistant Professor. Neurodevelopmental disorders, neuropsychiatric disease, neurodegenerative disease, immunology, pathogenesis.
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David Savage, Associate Professor. Synthetic biology and metabolism.
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David Schaffer, Professor. Neuroscience, biomolecular engineering, bioengineering, stem cell biology, gene therapy.
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Randy W. Schekman, Professor. Saccharomyces cerevisiae, organelle assembly, intracellular protein transport, assembly of cellular organelles, Alzheimer's Disease.
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Kristin Scott, Professor. Nerve cell connectivity in developing nervous systems, taste perception in the fruit fly, taste neural circuits, sensory maps in the brain.
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Bill Sha, Associate Professor. B cell gene regulation, fate determination, gene regulatory pathways, antibody-secreting plasma cells, memory B cells, apoptotic cells, B7 costimulatory ligands.
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Sarah Stanley, Assistant Professor. Mechanisms of pathogenesis and immune subversion in tuberculosis, protective immunity to tuberculosis, metabolic interactions between hosts and pathogens, development of novel therapeutics for tuberculosis, scientific capacity building, tuberculosis.
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Jeremy W. Thorner, Professor. Biochemistry, molecular genetics, cell biology, signal transduction mechanisms, protein kinase function and regulation, GPCRs, membrane biology, control of cell growth/morphology and division, regulation of gene expression by extracellular stimuli.
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Denis Titov, Assistant Adjunct Professor.
Robert T. Tjian, Professor. Eukaryotic molecular biology, biochemistry, cellular differentiation, chromatin function, RNA synthesis, single cell imaging, single molecule imaging.
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Elcin Unal, Assistant Professor. Gametogenesis, genetics, genomics and development.
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Russell E. Vance, Professor. Immunology, microbiology, infectious disease, molecular and cell biology.
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David A. Weisblat, Professor. Annelid developmental biology, leech embryo, evolution and development, cell fate determination, lineage tracingt.
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Matthew D. Welch, Professor. Biology, cell motility, the role of the actin cytoskeleton in cell locomotion, shape change, actin filament assembly, bacterial and viral pathogens.
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Astar Winoto, Professor. Cancer, genomics, apoptosis, innate immunity and infectious diseases, cell cycle, signal transduction, immune tolerance.
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Ahmet Yildiz, Assistant Professor. Single molecule biophysics, molecular motors, telomeres.
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Qiang Zhou, Professor. Biochemistry of HIV gene expression, transcriptional elongation, Tat activation, stage of transcriptional elongation, HIV replication, anti-HIV therapy.
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Roberto Zoncu, Assistant Professor. Biochemistry, biophysics, Structural Biology.
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Lecturers
Robin W. Ball, Lecturer.
P. Robert Beatty, Lecturer.
Natalia Caporale, Lecturer.
Isabelle Le Blanc, Lecturer.
Helen Lew, Lecturer.
David E. Presti, Senior Lecturer SOE.
Steve Takata, Lecturer.
Gary Joseph Wedemayer, Lecturer.
Visiting Faculty
Tamira M. Elul, Visiting Associate Professor.
Emeritus Faculty
Mark Alper, Professor Emeritus.
Bruce N. Ames, Professor Emeritus. Molecular biology, cancer, aging, mitochondrial decay, oxidants and antioxidants, micronutrient deficiencies and DNA damage, chronic inflammation and cancer.
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Giovanna F.-L. Ames, Professor Emeritus.
Clinton E. Ballou, Professor Emeritus.
Steven K. Beckendorf, Professor Emeritus. Genetics, biology, organogenesis, Drosophila, salivary morphogenesis.
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David R. Bentley, Professor Emeritus.
Phyllis B. Blair, Professor Emeritus.
Beth Burnside, Professor Emeritus. Cell biology of photoreceptors, cytoskeletal motors, morphogenetic events, photomembrane turnover.
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Richard Calendar, Professor Emeritus. Listeria monocytogenes, phage-based integration vector, Bacillus anthracis, vaccine.
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W. Zacheus Cande, Professor Emeritus. Genetics, cell biology, microbial biology, plant biology.
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M. J. Chamberlin, Professor Emeritus.
Alvin J. Clark, Professor Emeritus.
Thomas W. Cline, Professor Emeritus. Drosophila melanogaster, developmental genetics, sex determination.
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R. David Cole, Professor Emeritus.
John Gerhart, Professor Emeritus. Developmental biology, Xenopus laevis, Spemann's organizer, cortical rotation, cell cycle after fertilization, vegetal materials, blastula stage, egg cytoplasm.
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Robert M. Glaeser, Professor Emeritus.
Alexander N. Glazer, Professor Emeritus. Photosynthetic systems, phycobiliproteins, design of fluorescent probes, protein structure-function relationships, macromolecular complexes, environmental sciences, natural resource management.
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Stuart M. Linn, Professor Emeritus. Biology, enzymology of DNA metabolism, DNA repair and replication in mammalian cells, mechanisms of DNA damage by reactive oxygen species, structure of iron: DNA complexes, DNA damage and repair, DNA polymerases.
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Robert I. Macey, Professor Emeritus.
Terry Machen, Professor Emeritus. Physiology pathophysiology secretory epithelial cells, airway, ion transport, cell regulationm, imaging microscopy, calcium pH redox, electrophysiology, green fluorescent protein, genetic targeting, innate immune defense.
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G. Steven Martin, Professor Emeritus. Cell biology, signal transduction pathways, tumor virology, cell division cycle, viral and cellular oncoproteins, breast cancer.
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Howard C. Mel, Professor Emeritus.
Hsiao-Ping H. Moore, Professor Emeritus.
Satyabrata Nandi, Professor Emeritus.
Alexander V. Nichols, Professor Emeritus.
Hiroshi Nikaido, Professor Emeritus. Membrane biochemistry, bacterial physiology, bacteria, channel-forming proteins of the outer membrane, the diffusion of lipophilic compounds, mechanism and regulation of multidrug efflux transport systems, mycobacterial cell wall.
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W. Geoffrey Owen, Professor Emeritus. Biology, nervous system, membrane biophysics, retinal neurophysiology.
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Edward E. Penhoet, Professor Emeritus. Public health, health policy and management.
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Mu-Ming Poo, Professor Emeritus. Neurobiology, cellular and molecular mechanisms, axon guidance, synapse formation, activity-dependent refinement of neural circuits.
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Gerald M. Rubin, Professor Emeritus. Molecular genetics, molecular neurobiology, mapping and sequencing of the drosophila genome, genome organization and function, development and evolution.
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Harry Rubin, Professor Emeritus. Tumor biology, cell biology, regulation of neoplastic development, epithelial cells, oncogenic mutations, tumor development, RNA and DNA tumor viruses.
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Howard K. Schachman, Professor Emeritus. Physical biochemistry, biological macromolecules, aspartate transcarbamylase, revisiting allostery, holoenzyme, mutations, polypeptide chains, helical regions.
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+ Nilabh Shastri, Professor Emeritus. Cancer cells, mechanims of immunesurveillance, microbial pathogens, antigen genes, autoimmunity.
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Herbert H. Srebnik, Professor Emeritus.
Frank S. Werblin, Professor Emeritus. Retina, biological image processing, visual neuroscience.
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Gerald Westheimer, Professor Emeritus. Neurobiology, psychophysics, primate visual cortex, neural circuits, brain mechanisms, response modifications, active perception, learning, stereoscopic vision, optometryoptics of the eye, ophthalmic instrumentation.
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Fred H. Wilt, Professor Emeritus. Molecular embryology, cell biology, the regulation of gene expression, of sea urchin embryos, blastomeres, endoskeletal spicule of the larva, glycoproteins, immunoelectron microscopy, fluorescent labeling.
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Leon Wofsy, Professor Emeritus.
Robert S. Zucker, Professor Emeritus. Synaptic transmission, cellular neurophysiology, synaptic biophysics, properties of neural circuits, photolysis, vital dyes of vesicle membrane, electrophysiological techniques, neuromodulator.
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Contact Information
Department of Molecular and Cell Biology
497 Life Sciences Addition
Phone: 510-642-2651
Head Graduate Advisor
David Bilder, PhD
299 Life Sciences Addition
Phone: 510-642-5252
Graduate Advising
Graduate Affairs Office
299 Life Sciences Addition
Phone: 510-642-5252