About the Program

The Neuroscience Graduate Program at UC Berkeley is a unique, diverse PhD training program that offers intensive, integrated training in multiple areas of neuroscience research.

The program involves more than 55 faculty from different campus departments, with expertise ranging from molecular and cellular neuroscience to developmental neuroscience, systems and computational neuroscience, and human cognitive neuroscience.

We provide a highly interdisciplinary, intellectually dynamic training environment of coursework, research training, and mentoring, within a strong research program that produces fundamental advances in knowledge and cutting-edge techniques.

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Admissions

Admission to the University

Minimum Requirements for Admission

The following minimum requirements apply to all graduate programs and will be verified by the Graduate Division:

1. A bachelor’s degree or recognized equivalent from an accredited institution;
2. A grade point average of B or better (3.0);
3. If the applicant comes from a country or political entity (e.g., Quebec) where English is not the official language, adequate proficiency in English to do graduate work, as evidenced by a TOEFL score of at least 90 on the iBT test, 570 on the paper-and-pencil test, or an IELTS Band score of at least 7 (note that individual programs may set higher levels for any of these); and
4. Sufficient undergraduate training to do graduate work in the given field.

Applicants Who Already Hold a Graduate Degree

The Graduate Council views academic degrees not as vocational training certificates, but as evidence of broad training in research methods, independent study, and articulation of learning. Therefore, applicants who already have academic graduate degrees should be able to pursue new subject matter at an advanced level without need to enroll in a related or similar graduate program.

Programs may consider students for an additional academic master’s or professional master’s degree only if the additional degree is in a distinctly different field.

Applicants admitted to a doctoral program that requires a master’s degree to be earned at Berkeley as a prerequisite (even though the applicant already has a master’s degree from another institution in the same or a closely allied field of study) will be permitted to undertake the second master’s degree, despite the overlap in field.

The Graduate Division will admit students for a second doctoral degree only if they meet the following guidelines:

1. Applicants with doctoral degrees may be admitted for an additional doctoral degree only if that degree program is in a general area of knowledge distinctly different from the field in which they earned their original degree. For example, a physics PhD could be admitted to a doctoral degree program in music or history; however, a student with a doctoral degree in mathematics would not be permitted to add a PhD in statistics.
2. Applicants who hold the PhD degree may be admitted to a professional doctorate or professional master’s degree program if there is no duplication of training involved.

Applicants may apply only to one single degree program or one concurrent degree program per admission cycle.

Required Documents for Applications

1. Transcripts: Applicants may upload unofficial transcripts with your application for the departmental initial review. If the applicant is admitted, then official transcripts of all college-level work will be required. Official transcripts must be in sealed envelopes as issued by the school(s) attended. If you have attended Berkeley, upload your unofficial transcript with your application for the departmental initial review. If you are admitted, an official transcript with evidence of degree conferral will not be required.
2. Letters of recommendation: Applicants may request online letters of recommendation through the online application system. Hard copies of recommendation letters must be sent directly to the program, not the Graduate Division.
3. Evidence of English language proficiency: All applicants from countries or political entities in which the official language is not English are required to submit official evidence of English language proficiency. This applies to applicants from Bangladesh, Burma, Nepal, India, Pakistan, Latin America, the Middle East, the People’s Republic of China, Taiwan, Japan, Korea, Southeast Asia, most European countries, and Quebec (Canada). However, applicants who, at the time of application, have already completed at least one year of full-time academic course work with grades of B or better at a US university may submit an official transcript from the US university to fulfill this requirement. The following courses will not fulfill this requirement:
   • courses in English as a Second Language,
   • courses conducted in a language other than English,
   • courses that will be completed after the application is submitted, and
   • courses of a non-academic nature.

If applicants have previously been denied admission to Berkeley on the basis of their English language proficiency, they must submit new test scores that meet the current minimum from one of the standardized tests.

Where to Apply

Visit the Berkeley Graduate Division application page. 

Admission to the Program

Applicants to the program should have a bachelor's degree in science from a four-year college and at least one year of laboratory experience. Applicants are required to submit Graduate Record Examination (GRE) General Test scores, and are strongly encouraged to submit one GRE Subject Test score (in biochemistry and cell biology, chemistry, psychology, biology, computer science, or physics).

Doctoral Degree Requirements

Normative Time Requirements

Normative Time to Advancement

Step I: Lab Rotations and Presentations

During the first year of graduate study, each neuroscience graduate student spends three 10-week periods performing research projects in different faculty laboratories. The goal is to expose students to different techniques and approaches in neuroscience and to provide training in experimental design, critical analysis of data, and presentation of research findings. Performance in rotations is evaluated and graded. Rotations also allow students to identify the laboratory in which their thesis research will be performed. During the spring semester, students formally present results from the laboratory rotations in a dedicated course, NEUROSC 290 (Neuroscience First Year Research), designed to train students in clear, effective presentation of scientific findings.

Step II: Qualifying Exam

Students complete an oral qualifying exam during the spring semester of Year 2. This exam is structured around two written proposals—one in the student’s proposed area of thesis research, and the other in an area of neuroscience outside the thesis topic. During the exam, a faculty committee tests the student’s knowledge of these areas and general neuroscience. Students must demonstrate the ability to recognize important research problems, propose relevant experimental approaches, and display comprehensive knowledge of relevant subjects. Students must pass the qualifying examination before advancing to doctoral candidacy.

Normative Time in Candidacy

Step III: Dissertation

Students undertake research for the PhD dissertation under a four-person committee in charge of their research and dissertation. Students do original research using a wide variety of cutting-edge neuroscience methods. The students then write the dissertation based on the results of this research. On completion of the research and approval of the dissertation by the committee, the students are awarded the doctorate.

Total Normative Time

Total normative time is 5.5 years.

Time to Advancement

Curriculum

One Graduate Course in Each of the Following Three Foundational Areas

Students can either take one graduate level course from each category, or three graduate level courses chosen form two areas, plus a selected advance undergraduate course from a third area. They are taken in years 1-2.

1. Cellular, Molecular and Developmental Neuroscience

Course List

Code	Title	Units
Choose one:
MCELLBI 231	Advanced Developmental and Stem Cell Biology	4
MCELLBI 236	Advanced Mammalian Physiology	5
MCELLBI 240	Advanced Genetic Analysis	4
NEUROSC C261	Cellular and Developmental Neurobiology	3

2. Systems and Computational Neuroscience 

Course List

Code	Title	Units
Choose one:
PSYCH 210B	Proseminar: Cognition, Brain, and Behavior	3
VIS SCI 265	Neural Computation	3
NEUROSC C262	Circuit and Systems Neurobiology	3

3. Cognition, Brain and  Behavior 

Two Graduate Elective Seminars or Courses 

Students must take two additional elective courses. These can be either graduate-level seminars or graduate-level lecture courses, and can be 1 unit or more. These are typically taken years three-four. You may also select a foundation course as an elective. Consult your thesis adviser and thesis committee to select the most appropriate course for you.

Course List

Code	Title	Units
Neuro-Related Seminar Courses
EL ENG 290A	Advanced Topics in Electrical Engineering: Advanced Topics in Computer-Aided Design	1-3
EL ENG 290B	Advanced Topics in Electrical Engineering: Advanced Topics in Solid State Devices	1-3
EL ENG 290C	Advanced Topics in Electrical Engineering: Advanced Topics in Circuit Design	1-3
EL ENG 290D	Advanced Topics in Electrical Engineering: Advanced Topics in Semiconductor Technology	1-3
EL ENG 290F	Advanced Topics in Electrical Engineering: Advanced Topics in Photonics	1-3
EL ENG 290N	Advanced Topics in Electrical Engineering: Advanced Topics in System Theory	1-3
EL ENG 290O	Advanced Topics in Electrical Engineering: Advanced Topics in Control	1-3
EL ENG 290P	Advanced Topics in Electrical Engineering: Advanced Topics in Bioelectronics	1-3
EL ENG 290Q	Advanced Topics in Electrical Engineering: Advanced Topics in Communication Networks	1-3
EL ENG 290S	Advanced Topics in Electrical Engineering: Advanced Topics in Communications and Information Theory	1-3
EL ENG 290T	Advanced Topics in Electrical Engineering: Advanced Topics in Signal Processing	1-3
EL ENG 290Y	Advanced Topics in Electrical Engineering: Organic Materials in Electronics	3
LINGUIS 290A	Topics in Linguistic Theory: Syntax	3
LINGUIS 290B	Topics in Linguistic Theory: Semantics	3
LINGUIS 290D	Topics in Linguistic Theory: Pragmatics	3
LINGUIS 290E	Topics in Linguistic Theory: Phonology	3
LINGUIS 290F	Topics in Linguistic Theory: Diachronic Linguistics	3
LINGUIS 290H	Topics in Linguistic Theory: Linguistic Reconstruction	3
LINGUIS 290L	Additional Seminar on Special Topics to Be Announced	3
LINGUIS 290M	Topics in Linguistic Theory: Psycholinguistics	3
LINGUIS 290R	Course Not Available	3
MCELLBI 290	Graduate Seminar	1
PSYCH 290B	Seminars: Biological	2
PSYCH 290E	Seminars: Perception	2
PSYCH 290H	Seminars: Developmental	2
PSYCH 290I	Seminars: Personality	2
PSYCH 290J	Seminars: Social	2
PSYCH 290K	Seminars: Clinical	2
PSYCH 290P	Seminars: Additional Seminars on Special Topics to Be Announced	2
PSYCH 290Q	Seminars: Cognition	2
PSYCH 290Z	Seminars	1-3
VIS SCI 298	Group Studies, Seminars, or Group Research	1-6
Statistics
PSYCH 102	Methods for Research in Psychological Sciences	3
PSYCH 205	Data Analysis	3
STAT 150	Stochastic Processes	3
STAT 151A & STAT 151B	Linear Modelling: Theory and Applications and Course Not Available	4
STAT 153	Introduction to Time Series	4
STAT 204	Probability for Applications	4
STAT C241A	Statistical Learning Theory	3
STAT C241B	Advanced Topics in Learning and Decision Making	3
STAT 248	Analysis of Time Series	4
Mathematics
MATH 118	Fourier Analysis, Wavelets, and Signal Processing	4
MATH 220	Introduction to Probabilistic Methods in Mathematics and the Sciences	4
Computer Science and Programming
COMPSCI C280	Computer Vision	3
Electrical Engineering
EL ENG 120	Signals and Systems	4
EL ENG 123	Digital Signal Processing	4
EL ENG 126	Probability and Random Processes	4
EL ENG 221A	Linear System Theory	4
EL ENG 225A	Digital Signal Processing	3
EL ENG 225B	Digital Image Processing	3
EL ENG 226A	Random Processes in Systems	4
EL ENG 229A	Information Theory and Coding	3
Bioengineering
BIO ENG C218	Stem Cells and Directed Organogenesis	3
BIO ENG C219	Protein Engineering	3
BIO ENG 231	Introduction to Computational Molecular and Cellular Biology	4
BIO ENG 243	Computational Methods in Biology	4
BIO ENG 263	Principles of Molecular and Cellular Biophotonics	4
BIO ENG C265	Principles of Magnetic Resonance Imaging	4
Vision Science
VIS SCI 212B	Course Not Available	2
VIS SCI 212E	Color Vision and Visual Sensitivity	2
VIS SCI 212F	Spatial and Binocular Vision, Eye Movements, and Motion Perception	2
VIS SCI 212G	Molecular Genetics of Vertebrate Eye Development and Diseases	2

Other Degree Requirements 

Course List

Code	Title	Units
Pedagogy course
NEUROSC 290	Neuroscience First Year Research	2
NEUROSC 294	Neuroscience Graduate Student Presentation Seminar	1
Students must also complete a 1-semester course in Applied Statistics in Neuroscience, or an equivalent approved course in statistics or quantitative analysis methods.

Time in Candidacy

Dissertation Presentation/Formal Exit Seminar

There is no formal defense of the completed dissertation. Neuroscience students are required to publicly present a thesis seminar about their dissertation research in their final year.

Required Professional Development

Presentations

During their fourth year of study, students are required to make a presentation on the progress of their thesis work while enrolling in NEUROSC 294 (Neuroscience Graduate Student Presentation Seminar), a journal club, for a letter grade.

Teaching

Neuroscience students are required to serve as graduate student instructors (GSIs) for two semesters. Whenever possible, GSI assignments are determined with an eye toward student research interests. Teaching occurs during fall semester of the second year and spring semester of the third. Teaching affords students supervised experience in a variety of educational situations, including labs, discussion sections, and demonstrations. GSIs also participate in record-keeping, grading, advising, and student consultations.

GSIs are evaluated by both supervising faculty and the students they teach. These evaluations become a permanent part of the student file. Deserving GSIs are nominated for the Outstanding Graduate Student Instructor Award.

Courses

Neuroscience

Faculty and Instructors

+ Indicates this faculty member is the recipient of the Distinguished Teaching Award.

Faculty

Hillel Adesnik, Assistant Professor.

Martin S. Banks, Professor. Stereopsis, virtual reality, optometry, multisensory interactions, self-motion perception, vision, depth perception, displays, picture perception, visual ergonomics.
Research Profile

Helen Bateup, Assistant Professor. Molecular and cellular neuroscience, neurodevelopmental disorders, autism, epilepsy.
Research Profile

Diana Bautista, Associate Professor. Ion channels, sensory physiology, chemosensation, touch, thermosensation, somatosensory system.
Research Profile

George Bentley, Associate Professor. Hormones and behavior, neuroendocrinology of reproduction.
Research Profile

Sonia Bishop, Assistant Professor.

Steve Brohawn, Assistant Professor.
Research Profile

Jose M. Carmena, Professor. Brain-machine interfaces, neural ensemble computation, neuroprosthetics, sensorimotor learning and control.
Research Profile

Christopher J. Chang, Professor. Chemistry, inorganic chemistry, neuroscience, bioinorganic chemistry, general physiology, organic chemistry, new chemical tools for biological imaging and proteomics, new metal complexes for energy catalysis and green chemistry, chemical biology.
Research Profile

Anne Collins, Assistant Professor. Human learning, decision-making and executive functions; Computational modeling at multiple levels (cognitive and neuroscience); Behavioral, EEG, drug and genes studies in healthy or patient populations.Human learning, decision-making and executive functions; Computational modeling at multiple levels (cognitive and neuroscience); Behavioral, EEG, drug and genes studies in healthy or patient populations.
Research Profile

Mark T. D'Esposito, Professor. Cognitive neuroscience, psychology, working memory, frontal lobe function, functional MRI, neurology, brain imaging, dopamine.
Research Profile

Yang Dan, Professor. Neuronal circuits, mammalian visual system, electrophysiological, psychophysical and computational techniques, visual cortical circuits, visual neurons.
Research Profile

Michael Deweese, Assistant Professor. Machine learning, computation, systems neuroscience, auditory cortex, neural coding.
Research Profile

Andrew Dillin, Professor.

+ Dan Feldman, Associate Professor. Neurobiology, learning, neurophysiology, sensory biology.
Research Profile

Marla B. Feller, Professor. Neurophysiology, developmental neuroscience.
Research Profile

John Gerard Flannery, Professor. Neurobiology, optometry, vision science, cell and molecular biology of the retina in normal and diseased states.
Research Profile

David Foster, Professor.

Darlene Francis, Associate Professor.

Jack L. Gallant, Professor. Vision science, form vision, attention, fMRI, computational neuroscience, natural scene perception, brain encoding, brain decoding.
Research Profile

Gian Garriga, Professor. Developmental neurobiology; molecular genetics, development of nervous systems, cell division, cell migration, axonal pathfinding, caenorhabditis elegans.
Research Profile

Tom Griffiths, Associate Professor. Machine learning, computational models of human cognition, Bayesian statistics, cultural evolution.
Research Profile

Ming Hsu, Assistant Professor. Cognitive neuroscience, experimental economics, behavioral economics, neuroeconomics.
Research Profile

Ehud Y. Isacoff, Professor. Ion channel function, synaptic plasticity, neural excitability, synaptic transmission, the synapse.
Research Profile

Rich Ivry, Professor. Cognitive neuroscience, behavior, cognition, brain, attention, coordination, psychology, motor and perceptual processes in normal and neurologically impaired populations, temporal processing, executive control.
Research Profile

Lucia F. Jacobs, Professor. Cognitive and brain evolution, adaptive patterns in spatial memory, spatial navigation, cognitive sex differences and decision making.
Research Profile

William J. Jagust, Professor. Neuroscience, cognition, brain aging, dementia, imaging, Alzheimer's disease.
Research Profile

Na Ji, Associate Professor. Biophysics.

Daniela Kaufer, Associate Professor. Neuroscience, stress, neural stem cells, epilepsy, traumatic brain injury, blood brain barrier, prosocial behavior.
Research Profile

Stanley A. Klein, Professor. Optometry, vision science, spatial vision modeling, psychophysical methods and vision test design, corneal topography and contact lens design, source localization of evoked potentials, fMRI, amblyopia.
Research Profile

Robert Thomas Knight, Professor. Cognitive neuroscience, language, physiology, memory, attention, psychology, working memory, neuropsychology, human prefrontal cortex, neural mechanisms of cognitive processing, sensory gating, sustained attention, ad novelty detection.
Research Profile

Richard H. Kramer, Professor. Cells, synaptic transmission, chemical signaling between neurons, ion channels, electrical signals, chemical reagents, synapses.
Research Profile

Lance Kriegsfeld, Associate Professor. NeuroendocrinologyCircadian Biology, Neuroimmunology, cancer biology, animal behavior.
Research Profile

Stephan Lammel, Assistant Professor. Neuroscience, Optogenetics, dopamine, addiction, depression.
Research Profile

Dennis M. Levi, Professor. Optometry, vision science, pattern vision, abnormal visual development.
Research Profile

Chunlei Liu, Professor.

Evan W. Miller, Assistant Professor.

John Ngai, Professor. Nervous system, molecular and cellular mechanisms of olfaction, detection of odors, odorant receptors, olfactory neurons, DNA microarray technologies, genome-wide patterns of gene expression.
Research Profile

Bruno Olshausen, Professor. Visual perception, computational neuroscience, computational vision.
Research Profile

Mu-Ming Poo, Professor. Neurobiology, cellular and molecular mechanisms, axon guidance, synapse formation, activity-dependent refinement of neural circuits.
Research Profile

Teresa Puthussery, Assistant Professor. Retinal Neurobiology and Neurophysiology.
Research Profile

David Schaffer, Professor. Neuroscience, biomolecular engineering, bioengineering, stem cell biology, gene therapy.
Research Profile

Kristin Scott, Professor. Nerve cell connectivity in developing nervous systems, taste perception in the fruit fly, taste neural circuits, sensory maps in the brain.
Research Profile

Arthur P. Shimamura, Professor. Cognitive neuroscience, behavior, cognition, brain, psychology, frontal lobe function, basic memory research.
Research Profile

Michael Silver, Associate Professor. Cognitive neuroscience, pharmacology, learning, attention, visual perception, neuroimaging.
Research Profile

Fritz SOMMER, Adjunct Professor. Bayesian methods, information theory, memory, sensory processing, visual system.
Research Profile

Mark A. Tanouye, Professor. Genetics, neuroanatomy, electrophysiology, mechanisms of nervous system structure and function, drosophila mutants.
Research Profile

W. Rowland Taylor, Professor. Retinal circuit function, neural architecture, immunohisochemical studies.
Research Profile

Frederic Theunissen, Professor. Behavior, cognition, brain, psychology, birdsong, vocal learning, audition, neurophysiology, speech perception, computational neuroscience, theoretical neuroscience.
Research Profile

Jonathan David Wallis, Professor. Prefrontal cortex, neurophysiology, executive control, decision making.
Research Profile

David Whitney, Professor. Cognitive neuroscience, cognition, attention, visual perception, vision, visually guided action.
Research Profile

Linda Wilbrecht, Assistant Professor. Neuroscience, addiction, early life adversity, adolescence.
Research Profile

Michael Yartsev, Assistant Professor. Neuroscience, engineering.
Research Profile